Education and Career

1968–1971 Leicester University, B.Sc. Biological Sciences.
Class 2i,Science Faculty Prize, 1970

1971–1974 Leicester University, Ph.D. Physiology,

1981–1986 The Open University, B.A. Mathematics.
Class: First

1991 London University, D.Sc. Physiology Regulation
of muscle contractility and excitability

1995–2008 Professor of Cell Physiology, Institute of Urology,
University College London

1994–1995 Reader in Cell Physiology, Institute of Urology &
Nephrology, University College London.

1991–1993 Reader in Cell Physiology, Division of Physiology,
UMDS, University of London.
Vice-chairman, Division of Physiology, UMDS,
London University

1987–1991 Senior Lecturer in Physiology, UMDS, London

1979–1987 Lecturer in Physiology. St Thomas’ Hospital
Medical School (later UMDS), London University.

1978–1979 British Heart Foundation Lecturer in Cardiac
Physiology. Cardiothoracic Institute, London.

1977–1978 Research Assistant. Cardiothoracic Institute, London.

1976–1977 Teaching Assistant. Physiologisches Institüt,
University of Bern, Switzerland.

1975–1976 Royal Society Fellow. Physiologisches Institüt,
University of Bern, Switzerland.

Other positions

2002– Royal College of Surgeons of England: Court of

1991–1992 STEP cardiovascular course

2001–2002 The Physiological Society: Chairman of Executive

1999–2001 Committee Secretary;

1995–1999 Meetings Secretary

1994–1995 member of committee
Section editor (journals)
Neurourology & Urodynamics (Cell Biochemistry)
Nephron (Physiology); LUTS
International Continence Society Advisory Board
– Chairman, section Cell Biology

2009– External examiner. BSc Biological Sciences

2006–2009 BSc physiology (Bristol)

2005–2008 MBBChir (Cambridge)

2000–2004 BSc physiology (Belfast)

1999–2003 Cardiff

1996–1999 Leeds

1995–1998 MSc Physiology Kings College

1984–2009 PhD, MD, MS degree examiner

1990–1993 Gordon Conference; Chair ;

1987–1990 vice-Chair
Magnesium in biochemical processes
University of Surrey appointments
Medical School standing committee


The Physiological Society, The American Physiological Society, International Continence
Society, British Society for Cardiovascular Research


Post-graduate students: PhD students 12;
MD/MS 23 (all successful);
MBBS, Medicine: all aspects of physiology;
BSc: 1st-3rd year physiology BSc modules (UCL):
Membrane biophysics (UMDS);
MRCS, FRCAnaesth Applied physiology for surgeons and anaesthetists